History of research on the Hungarian Oniscidea (Isopoda) and assess-ment of the fauna
Abstract
The first data on the composition of the fauna of terrestrial woodlice fauna and on the distribution of species in Hungary are from 1794 and then from 1926. After the initial taxonomic, faunistic and anatomical works, production biology data were published in the 1960s, and then ecological and ecotoxicological works appeared from the 1970s and 1980s. In addition to the earlier ones, in the 21st century, results related to the functional role of the group, its practical applicability, and then behavioral and evolutionary biology research came to the fore in research. The country's area is very unevenly populated with species, with most of our data comes from the Transdanubian region. The current number of known species is 60, which can be divided into different types zoogeographically. The most common species ('big five'): Armadillidium vulgare, Hyloniscus riparius, Porcellium collicola, Trachelipus rathkii, Protracheoniscus politus. The rarest are the habitat specialist species with a narrow distribution, living in near-natural ecosystems. The basic fauna consists of European and cosmopolitan species with a wide ecological tolerance, while the endemic and rare species include Alpine, Illyrian, Balkan, Carpathian fauna elements, as well as introduced species mainly of tropical and Mediterranean origin. Our endemic species, Haplophthalmus hungaricus, was found in the north-eastern part of the country, while the special, endogeic Mesoniscus graniger is a member of the troglofauna, which was first described from the Baradla cave system. The species can also be grouped according to habitat type, altitude, and the degree of disturbance of the habitat, i.e. its naturalness, and based on this, both the species and their habitats can be classified.
References
Ábrahám, A. & Wolsky, A. (1929). Über die Nerven und Nervenendigungen der Antennen einiger Landasseln. Zoologischer Anzeiger, 84, 316–323.
Ábrahám, A. & Wolsky, A. (1930a). Über ein neues Sinnesorgan der Landisopoden. Zoologischer An-zeiger, 87, 87–93.
Ábrahám, A. & Wolsky, A. (1930b) Die Geruchsorgane der Landisopoden. Zeitschrift für Morpho-logie und Ökologie der Tiere, 17, 441–463. https://doi.org/10.1007/BF00419294
Balázs, G., Biró, A., Fišer, Ž., Fišer, C. & Herczeg, G. (2021). Parallel morphological evolution and habitat‐dependent sexual dimorphism in cave‐vs. surface populations of the Asellus aquaticus (Crustacea: Isopoda: Asellidae) species complex. Ecology and Evolution, 11(21), 15389–15403. https://doi.org/10.1002/ece3.8233
Bardgett, R. (2005). The biology of soil: a community and ecosystem approach. Oxford University Press.
Berisha, H., Horváth, G., Fišer, Ž., Balázs, G., Fišer, C. & Herczeg, G. (2023). Sex-dependent increase of movement activity in the freshwater isopod Asellus aquaticus following adaptation to a predator-free cave habitat. Current Zoology, 69(4), 418–425. https://doi.org/10.1093/cz/zoac063
Bölöni, J., Molnár, Z. & Kun, A. (szerk.) (2011). Magyarország Élőhelyei. Vegetációtípusok leírása és határozója, ÁNÉR 2011., MTA Ökológiai és Botanikai Kutatóintézet.
Campos-Filho, I. S., Araujo, P. B., Bichuette, M. E., Trajano, E. & Taiti, S. (2014). Terrestrial isopods (Crustacea: Isopoda: Oniscidea) from Brazilian caves. Zoological Journal of the Linnean Society, 172(2), 360–425. https://doi.org/10.1111/zoj.12172
Campos-Filho, I. S., Bichuette, M. E. & Taiti, S (2016). Three new species of terrestrial isopods Crus-tacea, Isopoda, Oniscidea) from Brazilian caves. Nauplius, 24(00), e2016001. https://doi.org/10.1590/2358-2936e2016001
Campos-Filho, I. S., Bichuette, M. E., Araujo, P. B. & Taiti, S. (2017). Description of a new species of Cylindroniscus Arcangeli, 1929 (Isopoda: Oniscidea) from Brazil, with considerations on the fa-mily placement of the genus. North-western journal of zoology, 13(2), 227–233.
Cloudsley-Thompson, J. L. (1988). Evolution and Adaptation of Terrestrial Arthropods, Springer. https://doi.org/10.1007/978-3-642-61360-9
Compére, P. (1991). Fine strukture and elaboration of the epicuticule and the pore canal system in tergi-te cuticule of the land isopod Oniscus asellus during a moulting cycle. In Juchault, P. & Mocquard, J. P. (eds.):’Third Symposium on the Biology of Terrestrial Isopods’ (pp. 169–175). Universiteit de Poitiers.
Csicsek, R. (2019). Közönséges gömbászka (Isopoda: Armadillidium vulgare) védekező viselkedésé-nek vizsgálata. http://hdl.handle.net/2437/266825
Csicsek, R. (2024). Urbanizációs változások hatása ízeltlábúak viselkedésére (PhD disszertáció, Debre-ceni Egyetem). https://dea.lib.unideb.hu/server/api/core/bitstreams/2b9c9cfa-92cf-4401-8acf-f5738dbc0b47/content
Csiki, E. (1926). Magyarország szárazföldi Isopodái (Isopoda terrestria Hungariae). Annales Historico-naturales Musei Nationales Hungarici, 23, 1–79.
Csonka, D., Halasy, K., Mrak, P., Strus, J. & Hornung, E. (2011). Eco-morphological comparison of three Armadillidium species (Crustacea: Oniscidea). In Zidar, P. & Strus, J. (eds): Proceedings of the 8th International Symposium of Terrestrial Isopod Biology, Bled, 2011.06.19–2011.06.23. Ljubljana. (pp. 45–46.)
Csonka, D., Halasy, K., Mrak, P., Štrus, J. & Hornung, E. (2012). Armadillidium-fajok (Isopoda: Oniscidea) élőhelyi adaptációjának morfológiai háttere. Természetvédelmi Közlemények, 18, 115–126. http://www.mbtktv.mtesz.hu/tvk2012.html
Csonka, D., Halasy, K. & Hornung, E. (2015). Histological studies on the marsupium of two terrestrial isopods (Crustacea, Isopoda, Oniscidea). ZooKeys, 515, 81–92. https://doi.org/10.3897/zookeys.515.9401
Csonka, D., Halasy, K., Buczkó, K. & Hornung, E. (2018). Morphological traits – desiccation toleran-ce – habitat characteristics: a possible key for distribution in woodlice (Isopoda: Oniscidea). ZooKeys, 801, 481–499. https://doi.org/10.3897/zookeys.801.23088
Csordás, B., Kontschán, J. & Hegyessy, G. (2005). Két ritka ászkarák (Oniscus asellus Linnaeus, 1758 és Protracheoniscus major (Dollfus, 1903)) újabb előfordulásai Magyarországon (Crustacea: Isopoda: Oniscidea). Folia historico naturalia Musei Matraensis, 29, 81–83.
Dailey, T. M., Claussen, D. L., Ladd, G. B. & Buckner, S. T. 2009. The effects of temperature, desicca-tion, and body mass on the locomotion of the terrestrial isopod, Porcellio laevis. Comparative Bioc-hemistry and Physiology, Part A 153, 162–166. https://doi.org/10.1016/j.cbpa.2009.02.005
Derbák, D., Dányi, L. & Hornung, E. (2018). Life history characteristics of a cave isopod (Mesoniscus graniger Friv.). ZooKeys, 801, 359–370. https://doi.org/10.3897/zookeys.801.23391
Dudich, E. (1942). Nachträge und Berichtigungen zum Crustaceen-Teil des ungarischen Faunen- kata-loges II. Fragmenta Faunistica Hungarica, 5, 1–13.
Edney, E. B. (1954). Woodlice and the land habitat. Biological Reviews (Camb), 29, 185–219. https://doi.org/10.1111/j.1469-185X.1954.tb00595.x
Edney, E. B. (1968). Transition from Water to Land in Isopod Crustaceans. American Zoologist, 8(3), 309–326. https://doi.org/10.1093/icb/8.3.309
ESDAC. European Soil Data Center. https://esdac.jrc.ec.europa.eu/ (utolsó megtekintés: 2025.07.31.)
Farkas, S. (1998). Population dynamics, spatial distribution, and sex ratio of Trachelipus rathkei Brandt (Isopoda: Oniscidea) in a wetland forest by the Drava river. Israel Journal of Zoology, 44, 323–331.
Farkas, S. (2004a). Data to the knowledge of the terrestrial isopod (Isopoda: Oniscidea) fauna of So-mogy county (Hungary: South Transdanubia). Somogyi Múzeumok Közleményei, 16, 313–323.
Farkas, S. (2004b). Data to the knowledge of the terrestrial Isopod (Isopoda: Oniscidea) fauna of the Mecsek Mountains (Hungary: South Transdanubia). Folia Comloensis, 13, 69–78.
Farkas, S. (2005). Data to the knowledge of the terrestrial Isopod (Isopoda: Oniscidea) fauna of Bara-nya county (Hungary: South Transdanubia). Acta Agraria Kaposváriensis, 9(1), 67–86.
Farkas, S. (2006). Tolna megye szárazföldi ászkaráfaunájának (Isopoda: Oniscidea) alapvetése. Állatta-ni Közlemények, 91(1), 29–42.
Farkas, S. (2007). The terrestrial isopod fauna of South Transdanubia (Hungary). Somogyi Múzeumok Közleményei B Természettudomány, 17, 159–168.
Farkas, S., Hornung, E. & Fischer, E. (1996). Toxicity of copper to Porcellio scaber Latr. (Isopoda) under different nutritional status. Bulletin of Environmental Contamination and Toxicology, 57(4), 582–588. https://doi.org/10.1007/s001289900230
Farkas, S. & Vilisics, F. (2006). A Mecsek szárazföldi ászkarák együttesei (Isopoda: Oniscidea). In Fazekas, I. (szerk.): A Mecsek állatvilága 1. / The fauna of the Mecsek Mts. 1, Hungary. Folia comloensis, 15, 25–34.
Farkas, S. & Vilisics, F. (2013). Magyarország szárazföldi ászkarák faunájának határozója (Isopoda: Oniscidea). Natura Somogyiensis, 23, 89–124. https://doi.org/10.24394/NatSom.2013.23.89
Ferrara, F., Paoli, P. & Taiti, S. (1991). Morphology of the pleopodal lungs in the Eubelidae (Crustacea, Oniscidea). In Juchault, P. & Mocquard, J. P. (eds): The Biology of Terrestrial Isopods. III. Pro-ceedings of the third International Symposium on the Biology of Terrestrial Isopods. Poitiers, France. 10&12 July, 1990. (pp. 169–175). Universiteit de Poitiers.
Ferrara, F., Paoli, P. & Taiti, S. 1997. An original respiratory structure in the xeric genus Periscyphis Gerstaecker, 1873 (Crustacea: Oniscidea: Eubelidae). Zooogischer Anzeiger, 235, 147–156.
Fischer, E., Majer, J. Hornung, E., Farkas, S. & Molnár, L. (1995). Sublethal Toxicity Test with the Woodlouse Porcellio scaber (Latr. 1814) (Isopoda: Porcellionidae): Progress report 1994 of SECOFASE (pp. 139–159).
Fischer, E., Farkas, S., Hornung, E. & Past, T. (1997). Sublethal effects of an organophosphorous insecticide, dimethoate, on the isopod Porcellio scaber Latr. Comparative Biochemistry and Physio-logy Part C: Pharmacology, Toxicology and Endocrinology, 116(2), 161–166. https://doi.org/10.1016/S0742-8413(96)00164-8
Forró, L. & Farkas, S. (1998). Checklist, preliminary distribution maps, and bibliography of woodlice in Hungary. Miscellanea Zoologica Hungarica, 12, 21–44.
Frivaldszky, J. (1865). Adatok a magyarhoni barlangok faunájához. Mathematikai és Természettudo-mányi Közlemények, 3, 2–52.
Gere, G. (1956). The examination of the feeding biology and the humificative function of Diplopoda and Isopoda. Acta Biologica Academiae Scientiarum Hungaricae, 6, 257–271.
Gere, G. (1964). Untersuchungen über die Temperaturtoleranz von Mesoniscus graniger J. Friv. (Crust. Isop.). Annales Universitatis Scientiarum Budapestinensis de Rolando Eötvös Nominatae Sectio Biologica, 7, 95–103.
Gere, G. (1965). Fütterungsversuche mit bodenbewohnenden Diplopoden und Isopoden in der Baradla-Höhle bei Aggtelek (Ungarn). Opuscula Zoologica, 5, 193–196.
Giurginca, A. (2006). On some Oniscidea and Diplopoda from Bucharest Romania. Archives of Bio-logical Sciences, 58, 31–35. https://doi.org/10.2298/ABS0601031G
Giurginca, A., Baba, S. C. & Munteanu, C-M. (2017). New data on the Oniscidea, Diplopoda and Chi-lopoda from urban parks of Bucharest. North-Western Journal of Zoology, 13(2), e161306 http://biozoojournals.ro/nwjz/content/v13n2/nwjz_e161306_Giurginca.pdf
Global Soil Partnership. Global Soil Biodiversity Initiative. https://www.globalsoilbiodiversity.org/ (utolsó megtekintés: 2025. 07.31).
Gongalsky, K. B., De Smedt, P., Kuznetsova, D. M., Segers, S., Gardini, P., Hornung, E., Korobush-kin, D. I., Orihuela-Rivero, R., Robla, J., Szlavecz, K., de Araujo, P. B., Barantal, S., Berg, M. P., Bichuette, M. E., Binder, S., Boeraeve, P., Burton, V. J., Byzov, P. S., Campos-Filho, I. S., ...Taiti, S. (2025). OniscidBase: A call for contributions to a global database to boost terrestrial isopod rese-arch. Soil Organisms, 97(3) (in press) https://doi.org/10.25674/466
Gruber, G. & Taiti, S. (2004). Morphology of the respiratory apparatus in the family Armadillidae (Crustacea, Isopoda, Oniscidea). 6th International Symposium on the Biology of Terrestrial Isopods. University of Aveiro. Aveiro, Portugal. Abstract Book (p. 63).
Gruner, H. (1966). Die Tierwelt Deutschlands. 53. Teil. Krebstiere oder Crustacea. V. Isopoda, 2. Lie-ferung, Jena. pp. 151–380.
Gruner, H.-E. & Tabacaru, I. G. (1963). Revision Der Familie Mesoniscidae Verhoeff, 1908 (Isopoda, Oniscoidea). Crustaceana, 6(1), 15–34. http://doi.org/10.1163/156854063X00318
Hanlon, R. D. G. & Anderson, J. M. (1980). Influence of macroarthropod feeding activities on mic-roflora in decomposing oak leaves. Soil Biology and Biochemistry, 12(3), 255–261. https://doi.org/10.1016/0038-0717(80)90071-1
Hassall, M., Turner, J. G. & Rands, M. R. W. 1987. Effects of terrestrial isopods on the decomposition of woodland leaf litter. Oecologia, 72, 597–604. https://doi.org/10.1007/BF00378988
Heeley, W. (1941). Observations on the life-histories of terrestrial isopods. Proceedings of the Zoologi-cal Society of London (B), 111, 79–149. https://doi.org/10.1111/j.1469-7998.1941.tb00044.x
Herczeg, G., Nyitrai, V., Balázs, G. & Horváth, G. (2022). Food preference and food type innovation of surface-vs. cave-dwelling waterlouse (Asellus aquaticus) after 60 000 years of isolation. Behavioral Ecology and Sociobiology, 76, 1–11. https://doi.org/10.1007/s00265-021-03109-x
Hoese, B. (1981). Morphologie und Funktion des Wasserleitungssystems der terrestrischen Isopoden (Crustacea, Isopoda, Oniscoidea). Zoomorphology, 98, 135–167. https://doi.org/10.1007/BF00310433
Hoese, B. (1982). Morphologie und Evolution der Lungen bei den terrestrischen Isopoden (Crustacea, Isopoda, Oniscoidea). Zoologische Jahrbücher, Abteilung für Anatomie und Ontogenie der Tiere, 107, 396–422.
Hoese, B. (1983). Struktur und Entwicklung der Lungen der Tylidae (Crustacea, Isopoda, Oniscoidea). Zoologische Jahrbücher, Abteilung für Anatomie und Ontogenie der Tiere, 109, 487–501.
Hoese, B. (1984). The marsupium in terrestrial isopods. In Sutton, S. L. & Holdich, D. (Eds.), Sympo-sia of the zoological Society of London, 53, 65–76.
Hoese, B. & Janssen, H. H. (1989). Morphological and physiological studies on the marsupium in ter-restrial isopods. Monitore Zoologico Italiano (N.S.) (Monograph), 4, 153–173.
Honek, A., Martinkova, Z., Saska, P. & Koprdova, S. (2009). Role of post-dispersal seed and seedling predation in establishment of dandelion (Taraxacum agg.) plants. Agriculture, ecosystems & environment, 134(1–2), 126–135. https://doi.org/10.1016/j.agee.2009.06.001
Hopkin, S. P. (1991). A key to the woodlice of Britain and Ireland, 7. Field Studies.
Hornung, E. (1981). Investigations on the productivity of the macrodecomposer isopod Trachelipus nodulosus C. L. Koch. Acta biologica Szeged, 27, 203–208.
Hornung, E. 1991. Isopod distribution in a heterogeneous grassland habitat. In Juchault, P. & Mocquard, J. P. (Eds.), Third Symposium on the Biology of Terrestrial Isopods (pp. 73–79). Uni-versiteit de Poitiers.
Hornung, E. (2011). Evolutionary adaptation of oniscidean isopods to terrestrial life: Structural – phy-siological – behavioural aspects. Terrestrial Arthropod Reviews, 4(2), 95–130. https://doi.org/10.1163/187498311X576262
Hornung, E. (2020). Skála - mintázat - élőhelyválasztás - életmenet: a szárazföldi ászkarákok (Isopo-da, Oniscidea) ökológiája. Akadémiai nagydoktori mű, pp. 155. https://real-d.mtak.hu/1255/ (utolsó megtekintés: 2025.08.01.)
Hornung, E. (2024). Terrestrial Adaptations of Crustaceans: The Challenges of Land Adaptations and Their Solutions in Terrestrial Isopods. In Saleuddin, S., Leys, S. P., Roer, R. D., Wilkie, I. C. (eds.) Frontiers in Invertebrate Physiology: A Collection of Reviews Volume 2: Crustacea. (pp. 327–387). Apple Academic Press. https://doi.org/10.1201/9781003405016
Hornung, E. (2025). Oniscidea bibliográfia 1998–2025. Állattani Közlemények, 110, in press.
Hornung, E., Fischer, E. & Farkas, S. (1998a). Isopod reproduction as a tool for sublethal-toxicity tests. Israel Journal of Zoology, 44(3–4), 445–450.
Hornung, E., Farkas, S. & Fischer, E. (1998b). Tests on the isopod Porcellio scaber. In Lokke, H., & Gestel, C. V. (Eds.), Handbook of soil invertebrate toxicity tests (pp. 207–226).Wiley.
Hornung, E., Tóthmérész, B., Magura, T. & Vilisics, F. (2007a). Changes of isopod assemblages along an urban-suburban-rural gradient in Hungary. European Journal of Soil Biology, 43(3), 158–165. https://doi.org/10.1016/j.ejsobi.2007.01.001
Hornung, E., Vilisics, F. & Szlávecz, K. (2007b). Szárazföldi ászkarák (Isopoda, Oniscidea) fajok tipi-zálása hazai előfordulási adatok alapján (különös tekintettel a sikeres megtelepedőkre). Természet-védelmi Közlemények, 13, 47–57.
Hornung, E., Vilisics, F. & Sólymos, P. (2008). Low alpha and high beta diversity in terrestrial isopod assemblages in the Transdanubian region of Hungary. In Zimmer, M., Cheikrouha, C. & Taiti, S. (Eds.), Proceedings of the International Symposium of Terrestrial Isopod Biology – ISTIB-7 (pp. 1–13). Shaker Verlag.
Hornung, E., Vilisics, F. & Sólymos, P. (2009). Ászkarák együttesek (Crustacea, Isopoda, Oniscidea) felhasználhatósága az élőhelyek természetességének minősítésében. Természetvédelmi Közlemé-nyek, 15, 381–395.
Hornung, E., Kásler, A. & Tóth, Z. (2018). The role of urban forest patches in maintaining isopod di-versity (Oniscidea). ZooKeys, 801, 371–388. https://doi.org/10.3897/zookeys.801.22829
Horváth, G., Garamszegi, L. Z., Bereczki, J., Urszán, T. J., Balázs, G. & Herczeg, G. (2019). Roll with the fear: environment and state dependence of pill bug (Armadillidium vulgare) personalities. The Science of Nature, 106(3), 7–11. https://doi.org/10.1007/s00114-019-1602-4
Horváth, G., Sztruhala, S. S., Balázs, G. & Herczeg, G. (2021). Population divergence in aggregation and sheltering behaviour in surface-versus cave-adapted Asellus aquaticus (Crustacea: Isopo-da). Biological Journal of the Linnean Society, 134(3), 667–678. https://doi.org/10.1093/biolinnean/blab093
Horváth, G., Kerekes, K., Nyitrai, V., Balazs, G., Berisha, H. & Herczeg, G. (2023). Exploratory beha-viour divergence between surface populations, cave colonists and a cave population in the water lo-use, Asellus aquaticus. Behavioral Ecology and Sociobiology, 77(1), 15. https://doi.org/10.1007/s00265-022-03288-1
Hudáková, J. & Mock, A. (2006). Suchozemské rovnakonőžky (Isopoda: Oniscidea) Pieninského národného parku. Entomofauna carpathica, 18, 47–55.
Hyžný, M. & Dávid, A. (2017). A remarkably well-preserved terrestrial isopod (Peracarida: Isopoda: Armadillidiidae) from the upper Oligocene of Hungary, with remarks on the oniscidean taphonomy. Paleontologia Electronica, 20.1.5A. https://doi.org/10.26879/615
IBP. International Biological Program. https://www.ebsco.com/research-starters/science/international-biological-program-ibp (utolsó megtekintés: 2025.07.31.)
Jones, A., Jeffery, S., Gardi, C., Montanarella, L., Marmo, L., Miko, L., Cranfield, K. R., Peres, G., Römbke, J. & van der Putten, W. (Eds.), 2004. European Atlas of Soil Biodiversity. Joint Research Centre’s Institute for Environment and Sustainability.
Kontschán, J. (2001a). A Bakonyvidék ászka (Crustacea: Isopoda: Oniscidea) faunájára új fajok. Folia Musei Historico-Naturalis Bakonyiensis, 18, 11–14.
Kontschán, J. (2001b). Két bakonyi telepített fenyves ászka együtteseinek (Crustacea: Ispoda: Onis-cidea) összehasonlító vizsgálata - Comparative study on the woodlice-fauna of two planted pine fo-rests. Folia Musei Historico-Naturalis Bakonyiensis, 18, 7–10.
Kontschán, J. (2001c). Adatok az Észak-Vértes és a Gerecse (Komárom-Esztergom megye) Peracarida (Crustacea: Isopoda et Amphipoda) faunájához. Komárom-Esztergom Megyei Múzeumok Közle-ményei, 8, 383–388.
Kontschán, J. (2001d). Adatok Majk (Észak-Vértes) magasabbrendű rák (Crustacea: Amphipoda et Isopoda et Decapoda) faunájához. Folia Historico-Naturalia Musei Matraensis, 25, 65–68.
Kontschán, J. (2002a). A Dunántúli-középhegység (Gerecse, Vértes és Bakony-vidék) Peracarida fau-nája. I. Vízi fajok (Amphipoda et Isopoda: Asellota) - Peracarida fauna of the Transdanubian Mountains (Gerecse, Vértes, and the region of Bakony). I. Aquatic species. Folia Musei Historico-Naturalis Bakonyiensis, 19, 25–30.
Kontschán, J. (2002b). The isopod and amphipod fauna of Fertő-Hanság National Park. In Mahunka S. (Ed.), The fauna of the Fertő-Hanság National Park (pp. 255–258). Magyar Természettudományi Múzeum.
Kontschán, J. (2002c). Ligidium fajok újabb adatai hazánkból és egy Magyarország faunájára új ászka-rák, a Ligidium intermedium Radu, 1950 előkerülése a Zempléni hegységből (Crustacea: Isopoda: Oniscidea). Folia entomologica hungarica, 63, 183–186.
Kontschán, J. (2003). Néhány ritka ászkarák (Crustacea: Isopoda: Oniscidea) újabb előfordulási adatai Magyarországról. Folia Historico-Naturalia Musei Matraensis, 27, 43–48.
Kontschán, J. & Berczik, Á. (2004). A Dunántúli-középhegység (Gerecse, Vértes, Bakony-vidék) Peracarida (Crustacea) faunája. II. Szárazföldi fajok (Isopoda: Oniscidea). Folia Musei Historico-Naturalis Bakonyiensis, 21, 73–82.
Kontschán, J. & Hornung, E. (2001). Peracarida (Crustacea: Isopoda et Amphipoda) fajok újabb adatai Magyarországról. In Isépy, I., Korsós, Z. & Pap, I. (szerk.): II. Kárpát-medencei Biológiai Szimpó-zium, Előadások összefoglalói (pp. 185–187). MBT & MTM.
Kontschán, J. (2004a). Magyarország faunájára új ászkarák (Reductoniscus constulatus Kesselyák, 1930 – Crustacea: Isopoda: Oniscidea) előkerülése az ELTE Füvészkertjéből (Budapest). Folia His-torico-Naturalia Musei Matraensis, 28, 89–90.
Kontschán, J. (2004b). Néhány adat az Északi-középhegység ászkarák faunájához (Crustacea: Isopoda: Oniscidea). Folia Historico-Naturalia Musei Matraensis, 28, 91–93.
Kontschán, J. (2004c). Reductoniscus costulatus Kesselyák, 1930, a new isopod species from Hungary. Folia Historico-Naturalia Musei Matraensis, 28, 89–90.
Kontschán, J. (2003). Néhány ritka ászkarák (Crustacea: Isopoda: Oniscidea) újabb előfordulási adatai Magyarországról. Folia Historico-Naturalia Musei Matraensis, 27, 43–48.
Kontschán, J., Hegyessy, G. & Csordás, B. (2006). Abaúj és Zemplén tájainak makroszkópikus rákjai (Crustacea). Abaúj-Zemplén Értékeiért Közhasznú Egyesület.
Korsós, Z., Hornung, E., Szlávecz, K. & Kontschán, J. (2002). Isopoda and Diplopoda of urban habi-tats: new data to the fauna of Budapest. Annales historico-naturales Musei nationalis hungarici, 94, 193–208.
Kula, H. & Larink, O. (1997). Development and standardization of test methods for the prediction of sublethal effects of chemicals on earthworms. Soil Biology and Biochemistry, 29(3–4), 635–639. https://doi.org/10.1016/S0038-0717(96)00030-2
Loksa, I. (1966). Die bodenzoologischen Verhältnisse der Flaumeichen-Buschwälder Südosteuropas. Akadémiai Kiadó.
Méhely, L. (1927). Gibt es zweiägige Trichonisciden? Zoologischer Anzeiger, 72, 1–4.
Méhely, L. (1929). Species generis Hyloniscus. Studia zoologica (Budapest), 1, 1–75.
Méhely, L. (1932). Adatok az ecsetcsápú rákok anatómiájához (Beiträge zur Anatomie der Trichonisci-den). Studia zoologica (Budapest), 2, 83–120.
Méhely, L. (1933). Csiki Ernő szárazföldi Isopodái című művének bírálata. Budapest: 1–18.
Mészárosné Póss, A., Südiné Fehér, A., Tóthné Bogdányi, F., & Tóth, F. (2021). The spread of the soil-borne pathogen Fusarium solani in stored potato can be controlled by terrestrial woodlice (Isopoda: Oniscidea). Agriculture, 12(1), 45. https://doi.org/10.3390/agriculture12010045
Mészárosné Póss, A., Tóthné Bogdányi, F. & Tóth, F. (2022). Consumption of fungi-infected fallen pear leaves by the common woodlouse. Acta Phytopathologica et Entomologica Hungarica, 57(1), 79–91.
Mock, A., Kania, G., Miklisová, D., Tajovský, K., Schlaghamerský, J. & Pižl, V. (2007). Terrestrial isopods (Oniscidea) of the Ojców National Park (Poland). Contributions to Soil Zoology in Central Europe II. (97–102). Institute of Soil Biology, Biology Centre, Academy of Sciences of the Czech Republic. .
Mödlinger, G. (1931). Beiträge zur Morphologie der Respirationsorgane der Landisopoden. Studia zoologica (Budapest), 2, 25–79.
Nagy, L. & Mizser, S. (2015). Az Armadillidium vulgare (Latreille, 1804) (Crustacea: Isopoda: Onis-cidea) toxikus elemtartalmának vizsgálata urbanizációs grádiens mentén. Természetvédelmi Közle-mények, 21, 186–193.
Navrátil, M. (2007). Stonožky, mnohonožky a suchozemští stejnonožci ve městě (Olomouc, Hodonín). MSc Thesis, Palacky University of Olomouc, 78 pp. http://myriapoda.upol.cz/tuf/pdf/papers/Navratil2007.pdf (utolsó megtekintés: 2025.08.11.)
Paoletti, M. G., Tsitsilas, A., Thomson, J., Taiti, S. & Umina, P. A. (2008). The flood bug, Australi-odillo bifrons (Isopoda: Armadillidae): A potential pest of cereals in Australia? Applied Soil Eco-logy, 39(1), 76–83. https://doi.org/10.1016/j.apsoil.2007.11.009
Paoli, P., Ferrara, F. & Taiti, S. (2002). Morphology and evolution of the respiratory apparatus in the family Eubelidae (Crustacea, Isopoda, Oniscidea). Journal of Morphology, 253, 272–289. https://doi.org/10.1002/jmor.10008
Papp, D., Simon, E., Nagy, L., Mizser, S. & Tóthmérész, B. (2019). The effect of urbanization on trace element concentration and symmetry of woodlice (Armadillidium vulgare Latreille, 1804). Biological Trace Element Research, 189(1), 251–258. https://doi.org/10.1007/s12011-018-1454-3
Paris, O. H. (1965). Vagility of P32 labelled isopods in grasslands. Ecology, 46(5), 635–648. https://doi.org/10.2307/1935003
Potočnik, F. (1979). Prispevek k poznavanju favne mokric (Isopoda terrestria) Slovenije. Biološki Vestník (Ljubljana), 27, 63–70.
Rabatin, S. C. & Stinner, B. R. (1988). Indirect effects of interactions between VAM fungi and soil-inhabiting invertebrates on plant processes. Agriculture, Ecosystems & Environment, 24(1–3), 135–146. https://doi.org/10.1016/0167-8809(88)90061-8
Rabatin, S. C. & Stinner, B. R. (1989). The Significance of Vesicular–Arbuscular Mycorrhizal Fungal–Soil Macroinvertebrate Interactions in Agroecosystems. Agricultural Ecology and Environment, 27(1–4), 195–204. https://doi.org/10.1016/B978-0-444-88610-1.50021-8
Riedel, P., Navrátil, M., Tuf, I. H. &. Tufová, J. (2009). Terrestrial isopods (Isopoda: Oniscidea) and millipedes (Diplopoda) of the city of Olomouc (Czech Republic). Contributions to Soil Zoology in Central Europe III (pp. 125–132). Institute of Soil Biology, Biology Centre, Academy of Sciences of the Czech Republic.
Saska, P. (2008). Granivory in terrestrial isopods. Ecological Entomology, 33(6), 742–747. https://doi.org/10.1111/j.1365-2311.2008.01026.x
Saska, P., Koprdova, S., Martinkova, Z. & Honěk, A. (2014). Comparing methods of weed seed expo-sure to predators. Annals of Applied Biology, 164(2), 301–312. https://doi.org/10.1111/aab.12102
Schmalfuss, H. (1975). Morphologie, Funktion und Evolution der Tergithöcker bei Landisopoden (Oniscoidea, Isopoda, Crustacea). Zeitschrift für Morphologie der Tiere, 80, 287–316. https://doi.org/10.1007/BF00302762
Schmalfuss, H. (1983). Asseln. Stuttgarter Beiträge zur Naturkunde, Serie C, 17, 1–28.
Schmalfuss, H. (2003). World catalog of terrestrial isopods (Isopoda: Oniscidea). Stuttgarter Beiträge zur Naturkunde, Serie A (Biologie), 654, 1–341.
Schmalfuss, H. (1984). Eco-morphological strategies in terrestrial isopods. In: Sutton, S. L & Holdich, D. (Eds.), Symposia of the zoological Society of London, 53, 49–63.
Schmalfuss, H. (2018). Research on the biology of terrestrial Isopods: A historical survey. Acta Zoo-logica Academiae Scientiarum Hungaricae, 64(3), 193–215. https://doi.org/10.17109/AZH.64.3.193.2018
Schmidt, C. & Wägele, J. W. (2001). Morphology and evolution of respiratory structures in the pleopod exopodites of terrestrial Isopoda (Crustacea, Isopoda, Oniscidea). Acta Zoologica (Stockholm), 82, 315–330. https://doi.org/10.1046/j.1463-6395.2001.00092.x
Schmölzer, K. (1965). Bestimmungsbücher zur Bodenfauna Europas. Ordnung Isopoda (Landasseln). Lieferung 4 und 5, Berlin.
Schmölzer, K. (1974). Catalogus Faunae Austriae, Teil 8e, Isopoda. Vienna.
Singer, C., Bello, N. M. & Snyder, B. A. (2012). Characterizing prevalence and ecological impact of non-native terrestrial isopods (Isopoda, Oniscidea) in tallgrass prarie. Crustaceana, 85(12–13), 1499–1511. https://doi.org/10.1163/15685403-00003126
Smigel, J. T. & Gibbs, A. G. (2008). Conglobation in the pill bug, Armadillidium vulgare, as a water conservation mechanism. Journal of Insect Science, 8(44), 1–9. https://doi.org/10.1673/031.008.4401
Stloukal, E. (2001). Databanka fauny Slovenska–Stav a nové prírastky databáz v roku 2001. Folia fau-nistica slovaca, 6, 145–168.
Sutton, S. L. (1972). Invertebrate types: Woodlice. Ginn.
Strouhal, H. (1958). Neue südostalpine Trichoniscus Arten aus Österreich (Isopoda, Oniscoidea). An-nalen des Nathistorischen Museums in Wien, 62, 283–295.
Szlávecz, K. (1992). The role of terrestrial isopods (Isopoda, Oniscidea) in the decomposition of aquatic macrophyte detritus of Lake Balaton, Hungary. Opuscula Zoologica, 25, 103–112.
Szlávecz, K. (1993). Needle litter consumption by two terrestrial isopods, Protracheoniscus amoenus (C.L. Koch) and Cylisticus convexus (de Geer). Pedobiologia, 37, 57–64. https://doi.org/10.1016/S0031-4056(24)00086-6
Szlávecz, K. (1995). Diversity and spatial community structure of terrestrial isopods (Isopoda, Onis-cidea) in a mosaic of plant assemblages. In Alikhan, M. A. (Ed.), Terrestrial Isopod Biology, Crus-tacean Issues, 9, 97–106.
Szlávecz, K. & Pobozsny, M. (1995). Coprophagy in isopods and diplopods: a case for interaction. An-nales Zoologici Fennica, 196, 124–128.
Szlavecz, K., Vilisics, F., Tóth, Z. & Hornung, E. (2018). Terrestrial isopods in urban environments: an overview. ZooKeys, 801, 97–126. https://doi.org/10.3897/zookeys.801.29580
Taiti, S., Paoli, P. & Ferrara, F. (1998). Morphology, biogeography, and ecology of the family Arma-dillidae (Crustacea, Oniscidea). Israel Journal of Zoology, 44(3–4), 291–301.
Tartally, A., Hornung, E. & Espadaler, E. (2004). The joint introduction of Platyarthrus schoblii (Isopoda: Oniscidea) and Lasius neglectus (Hymenoptera: Formicidae) into Hungary. Myrmecolo-gische Nachrichten, 6, 61–66.
Tóth, Z., Dombos, M. & Hornung, E. (2023). Urban soil quality deteriorates even with low heavy metal levels: An arthropod‐based multi‐indices approach. Ecological Applications, 33(4), e2848. https://doi.org/10.1002/eap.2848
Tuf, I. H. (2003). Development of the community structure of terrestrial isopods (Crustacea, Isopoda, Oniscidea) after a summer flood. In Sfenthourakis. S., de Araujo, P. B., Hornung, E. & Schmal-fuss, H. The biology of terrestrial isopods V. (Crustaceana Monographs, 2) (pp. 231– 242). Brill Academic Publisher. https://doi.org/10.1163/9789047412854_018
Tuf, I. H. & Jeřábková, E. (2008). Diurnal epigeic activity of terrestrial isopods (Isopoda: Oniscidea). In Zimmer, M., Charfi-Cheikhrouha, F. & Taiti, S. (Eds.), Proceedings of the international sympo-sium on terrestrial isopod biology: ISTIB-07 (pp. 167–172). Shaker Verlag.
Tuf, I. H. & Tufová, J. (2005). Communities of terrestrial isopods (Crustacea: Isopoda: Oniscidea). In: Epigeon of oak-hornbeam forests of SW Slovakia. Ekológia, 24, 113–123. Suppl. 2/2005.
Tuf, I. H. & Tufová, J. (2008). Proposal of ecological classification of centipede, millipede and terrest-rial isopod faunas for evaluation of habitat quality in Czech Republic. Časopis Slezského zemského muzea. Série A, 57, 37–44.
Vilisics, F. (2005). Új fajok és ritkaságok Magyarország terresztris ászkafaunájában (Isopoda: Onis-cidea, Crustacea). In Korsós Z. (szerk.): IV. Kárpát-medencei Biológiai Szimpózium, Budapest, 2009. okt. 17–19. (pp. 479–485). Magyar Biológiai Társaság.
Vilisics, F. (2007). New and rare species in the isopod fauna of Hungary (Crustacea, Isopoda, Onis-cidea): results of field surveys and revisions. Folia Historico-Naturalia Musei Matraensis, 31, 115–123.
Vilisics, F. & Farkas, S. (2004). Összehasonlító faunisztikai vizsgálat a dél-dunántúli Babarcszőlösi-pikkely ászkafaunáján (Isopoda, Oniscidea). Állattani Közemények, 89, 17–25.
Vilisics, F., Sólymos, P. & Hornung, E. (2007). Habitat features and associated terrestrial isopod speci-es: a sampling scheme and conservation implications. In Tajovský, K., Schlaghamerský, J. & Pižl, V. (Eds.), Contributions to Soil Zoology in Central Europe II. (pp. 97–102). Institute of Soil Bio-logy, Biology Centre, Academy of Sciences of the Czech Republic.
Vilisics, F. & Hornung, E. (2009). Urban areas as hot-spots for introduced and shelters for native isopod species. Urban Ecosystems, 12, 333–345. https://doi.org/10.1007/s11252-009-0097-8
Vilisics, F. & Hornung, E. (2010a). Újabb adatok Magyarország szárazföldi ászkarák (Crustacea, Isopoda, Oniscidea) faunájához. Állattani Közlemények, 95(1), 87–120.
Vilisics, F. & Hornung, E. (2010b). A magyar szárazföldi ászkarák (Isopoda, Oniscidea) fauna értéke-lése. In Kőrösi, Á. (szerk.): SZÜSZI Előadások és poszterek összefoglalói (p. 28).
Vilisics, F., Nagy, A., Sólymos, P., Farkas, R., Kemencei, Z., Páll-Gergely, B., Kisfali, M. & Hornung, E. (2008). Data on the terrestrial isopoda fauna of the Alsó-hegy, Aggtelek National Park, Hungary. Folia faunistica Slovaca, 13(4), 19–22.
Vilisics, F., Sólymos, P., Nagy, A., Farkas, R., Kemencei, Z. & Hornung, E. (2011). Small scale gradi-ent effects on isopods (Crustacea, Oniscidea) in karstic sink holes. Biologia (Bratislava), 65, 409–505. https://doi.org/10.2478/s11756-011-0042-1
Wilson, G. D. F. (2008). Global diversity of Isopod crustaceans (Crustacea; Isopoda) in freshwater. Hydrobiologia, 595, 231–240. https://doi.org/10.1007/s10750-007-9019-z
Vona-Túri, D. & Szmatona-Túri, T. (2012a). Adatok a Mátra-hegység ászkarák (Crustacea: Isopoda: Oniscidea) faunájához, különös tekintettel az út menti élőhelyekre. Természetvédelmi Közlemények, 18, 537–548.
Vona-Túri, D. & Szmatona-Túri, T. (2012b). Szárazföldi ászkarákok (Crustacea: Isopoda: Oniscidea) élőhely preferenciájának ökológiai háttere. 9. Magyar Ökológus Kongresszus (Keszthely, 2012. szeptember 5–7.) Előadások és poszterek összefoglalói (p. 110).
Vörös, J. (2014). Kétéltűek molekuláris taxonómiája a Kárpát-medencében. Magyar Tudomány, 175(4), 423–427.
Wallwork, J. A. (1970). Ecology of Soil Animals. New York, McGraw–Hill.
Warburg, M. R. (1968). Behavioral Adaptations of Terrestrial Isopods. Integrative and Comparative Biology, 8(3), 545–559. https://doi.org/10.1093/icb/8.3.545
Warburg, M. R. 1987. Isopods and their terrestrial environment. Advances in Ecological Research, 17, 187–242. https://doi.org/10.1016/S0065-2504(08)60246-9
Warburg, M. R. (1993). Evolutionary biology of land isopods. Springer-Verlag. https://doi.org/10.1007/978-3-662-21889-1
Wilson, G. D. F. (2008). Global diversity of Isopod crustaceans (Crustacea; Isopoda) in freshwater. Hydrobiologia, 595, 231–240. https://doi.org/10.1007/s10750-007-9019-z
WoRMS (2025). World Register of Marine Species. Isopoda. https://www.marinespecies.org/aphia.php?p=taxdetails&id=1131
